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Born in Gloucester, 1909; educated at the Crypt Grammar School, Gloucester, 1920-1928; graduated from St John's College Cambridge with a first class degree in both parts of the Natural Science Tripos (Part ll Biochemistry); began postgraduate research in the Biochemistry Department at Cambridge, receiving his PhD for 'Some comparative studies on phosphagen', 1934; principal research interest was comparative biochemistry; Fellow of St John's College Cambridge, 1936-1941; worked under Sir Frederick Gowland Hopkins as Demonstrator in Biochemistry, 1936-1943; also worked for periods at marine biological stations in France and at the Marine Biological Laboratory, Plymouth, in the 1930s; undertook a series of investigations of the pharmacology and physiology of Ascaris lumbricoides, 1940-1949; Lecturer in the Biochemistry Department at Cambridge, 1943-1950; Senior Fellow of the Lalor Foundation, USA, carrying out research into the phosphagen of the invertebrates at the Marine Biological Laboratories at Woods Hole, Massachusetts, 1948; as Joint Honorary Secretary and member of the Congress and Executive Committees, active in the organisation of the First International Congress of Biochemistry, in Cambridge, 1949; Professor of Biochemistry at University College London (UCL), 1950-1969; his reputation as an educator was one of the principal reasons for his appointment; established the first undergraduate biochemistry course at the College and orientated the biochemistry department as a branch of biological rather than chemical science; awarded the Cortina Ulisse Prize for the Italian edition of Dynamic Aspects of Biochemistry, 1952; after his move to UCL, his principal research interests were the comparative biochemistry of nitrogen metabolism and water shortage effects on the ureotelic metabolism; carried out research on ureogenesis in elasmobranch fishes during a period as Visiting Professor at the Scripps Institution of Oceanography, La Jolla, University of California, 1956-1957; author of several influential books on biochemistry; died, 1969. Publications include: An Introduction to Comparative Biochemistry (1937); Dynamic Aspects of Biochemistry (1947); The Nature of Biochemistry (1962).