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Born, 1872; educated King's College, Cambridge, 1893-1897; research at Cambridge Physiological Laboratory, 1897-; lecturer at King's College, Cambridge, 1899; junior demonstrator in the physiological laboratory, 1904; senior demonstrator, 1907; fellow of the Royal Society, 1910; assistant tutor King's College, Cambridge, 1910; expedition to Tenerife, 1910; expedition to Monte Rosa, 1911; CBE, 1918; reader King's College, Cambridge, 1919; high-altitude expedition to Cerro de Pasco in Peru, in order to study pulmonary gas exchange, blood biochemistry, and several other topics, 1921-1922; professor of physiology at Cambridge, 1925; Copley medal of the Royal Society, 1943; died, 1947.
Publications: The Respiratory Function of the Blood (1914, 2nd edition, 1925).