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Born in 1784, in Axminster, Devon, William Buckland had developed an interest in natural history and geology whilst exploring the local woods and quarries with his father Charles Buckland, the rector of Templeton and Trusham. Buckland was initially home schooled by his father, but in 1797 entered Blundell's School in Tiverton, enrolling the following year at St Mary's College, Winchester.
With the help of his uncle, John Buckland, he entered Corpus Christi College, Oxford, as a scholar on the Exeter Foundation, obtaining his BA in classics and theology three years later. Buckland was elected Fellow of the College in 1808 when he was ordained. During this time he had been attending the lectures of John Kidd, reader in mineralogy at Oxford and when Kidd resigned in 1813, Buckland succeeded him. That same year he became a Member of the Geological Society. In 1818 he was appointed to the new readership in geology and his inaugural lecture of 1819 'Vindiciae geologicae', compiled with the help of his mentor the Rev William Daniel Conybeare, explained that the facts of geology were conversant with the record of the creation of the Earth found in the Bible - notably the presence of older gravels (diluvium) which could not be explained through normal river deposits and therefore provided evidence of a 'universal deluge', that is Noah's flood.
Buckland's most important early geological work was on fossil cave faunas, principally his excavations of the Kirkdale Cavern, Yorkshire. There he discovered the bones of hyenas and other exotic animals such as an elephant, hippomatus and rhinocerus. His theory that the cave was a hyena den and the exotic animals had been dragged in as their prey was backed up by his experiments on and observation of modern hyenas. The results were published as 'Reliquiae diluvianae' (1823), and his discoveries at Kirkdale won him the Royal Society's Copley medal in 1822.
Due to his modest income from his two readerships and teaching, by 1825 Buckland was considering leaving Oxford for a more lucrative position, such as vicar, elswhere. However with the help of influential friends such as Sir Robert Peel, Buckland managed to obtain the position of canon to Christ Church which had an income of five times his previous salary. In December of that year he married Mary Morland, who had been helping him with his work and also illustrating his papers for a number of years.
In the field, Buckland notoriously dressed in a rather eccentric manner, always wearing his academic gown and carrying a large blue bag from which he would draw out his latest finds such as fossil faeces of giant marine reptiles. Buckland had found and identified these 'Coprolites', the term he coined for fossil faeces, in Lyme Regis when he worked with the fossil collector Mary Anning.
Buckland continued to research and publish over the next two decades, notably his influential treatise 'Geology and Mineralogy' of 1836, in which he abandoned his former belief in the geological effects deriving from the biblical flood. Indeed after a visit to Switzerland to see his friend Louis Agassiz in 1838, the Swiss naturalist had convinced him that glaciation had been more extensive in the past - leading Buckland to reinterpret his early theories and observations of a universal flood as evidence instead for the new glacial theory of an ice age. Buckland was extremely active within the Geological Society, serving twice as its President between 1824-1826 and 1839-1841, and winning the Wollaston Medal (the highest award bestowed by the Society) in 1848.
In 1845, Buckland had become dissatisfied with academic life in Oxford and accepted the appointment of dean of Westminster, coupled with the rectorship of Islip, near Oxford, although he still continued to lecture on geology in the university town. However by 1850 his diminishing mental health, possibly resulting from a fall from a coach a few years earlier, prevented Buckland from performing his duties as dean or professor. He retired to Islip but was later placed in The Retreat, a mental asylum in Clapham where he died on 14 August 1856.