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Born, 1899; Educated Perth Academy; University of St. Andrews; Assistant Medical Officer, Murray Royal, Perth, 1923; Lecturer in Epidemiology, 1924; Assistant in Bacteriology, St Andrews. Worked with Professors WJ Tulloch and WL Burgess; produced two MRC monographs with them, 1927; Research Associate, Connaught Laboratories, Toronto, 1931; Lecturer in Epidemiology, School of Hygiene, University of Toronto, 1932; Secretary of School of Hygiene, 1935-1945; Member of joint US-Canadian Commission on cattle industry and pests. Helped to write Memoranda for the National Research Council of Canada, 1942-1946; Associate Professor of Virus Infections, School of Hygiene, 1946; Elected President of the American Society of Bacteriologists and awarded US Typhus Commission Medal, 1946; Returned to England to work on cancer tumours at Imperial Cancer Research Fund Laboratory, Mill Hill, 1947; In charge of laboratory, Mill Hill, 1949-1957; retired, 1964; died, 1978.