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Born, 1909; work in the Department of Biochemistry, University College London, 1932-1933; Research in the Department of Physiology, University College, 1933-1936; Travelling Fellow of the Rockefeller Foundation, University of Pennsylvania Medical School, 1936-1937; Boit Memorial Fellow in Medical Sciences, Department of Physiology, University College, 1937-1939; Associate Professor of Physiology, Dalhousie University, Halifax, Nova Scotia, 1939-1942; Senior Experimental Officer, Ministry of Supply, 1942-1946; with James Danielli published pioneering work on the structure of cell membranes, 1943; Staff of Medical Research Council in collaboration with University College, 1946-1976; Research Fellow in the Department of Physiology, King's College London, 1976-1996; died, 1996. Publications: Co-authored with James Danielli, The permeability of natural membranes (Cambridge, 1943); The physiology of the eye (London, 1949); A textbook of general physiology (London, 1951); Physiology of the ocular and cerebrospinal fluids (London, 1956); Physiology of the cerebrospinal fluid (London, 1967); Co-authored with Malcolm Segal, Introduction to physiology (London, 1975-1980); Co-authored with Keasley Welch and Malcolm Segal, Physiology and pathophysiology of the cerebrospinal fluid (Edinburgh, 1987); An introduction to the blood-brain barrier (Basingstoke, 1993); edited The eye (New York, 1969-1977). He also published numerous articles in learned journals.