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Born 1865; educated Manchester Grammar School and Owens College, Manchester; Assistant in the Zoological Department, British Museum (Natural History), London, 1886-1887; Demonstrator and Assistant Lecturer in Biology, University of Melbourne, Australia, 1888-1894; Professor of Biology, Canterbury College, University of New Zealand, 1894-1903; Professor of Zoology, South African College, Cape Town, South Africa, 1903-1905; Chair of Zoology, King's College London, 1905-1925; member of the Royal Society's Committee for the Investigation of Grain Pests, 1917-1919; died 1925.
Publications: A monograph of the Victorian sponges (Melbourne, 1891); editor of Animal life and human progress (Constable and Co, London, 1919); Outlines of evolutionary biology (Constable and Co, London, 1912); Porifera. Part I. Non-Antartic sponges (London, 1924); editor Problems of modern science (George G Harrap and Co, London, 1922); The Anatomy of an Australian Land Planarian; The biological foundations of society (Constable and Co, London, 1924); An introduction to the study of Botany (Melville, Mullen and Slade, Melbourne and London, 1892).