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Born, 1900; Educated at Wychwood School Oxford; Madras College, St Andrews, 1916; Edinburgh University to read Zoology, 1918; B.Sc: works with Dr F A E Crew; goes to Cambridge as Scientific Assistant to Dr T S P Strangeways with MRC funding, 1923; Junior Beit Fellowship, Ph.D, 1924; Senior Beit Fellow; Director of the Strangeways Research Laboratory, 1929-1970; Messel Research Fellow of the Royal Society, 1931-1943; D.Sc Edinburgh, 1932; Foulerton Research Fellow of the Royal Society, 1943-1967; FRS, 1952; DBE, Royal Society Research Professor, 1963; retires as Director of the Strangeways Laboratory, 1970; Research Worker in Division of Immunology, Department of Pathology, University of Cambridge, 1970-1979; Research worker at Strangeways Laboratory, 1979-1986; died, 1986.