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Born 1908; educated, King's College School, Wimbledon, Queens' College Cambridge; BA 1930, 1st Class Hons Natural Science Tripos Pt II, 1931; Assistant in Zoology, University of Glasgow, 1932-1937; MA 1934; Assistant Lecturer and Lecturer in Zoology, University College, Cardiff, 1937-1948; MSc (Wales) 1943; Professor of Biology and Head of Biology Department, Guy's Hospital Medical School, 1948-1972; Reader in Biology, University of London, 1948-1955; Professor of Biology, University of London, 1955-1972; Chairman, British Universal Film Council, 1959-1963, 1967-1969; Emeritus Professor, University of London, 1972-1979; Fellow Cambridge Philosophical Society; Fellow Linnean Society; Fellow, Institute of Biology; Fellow, Zoological Society; died, 1979.
Publications: Report on Stomatopod Larvae, Cumacea and Cladocera [1930]; Stomatopod Larvæ, etc (London, 1939); various scientific papers, mainly dealing with the comparative study of the heart and blood system of vertebrate animals.