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Born 1903; educated at Imperial College, 1920-1923; Beit Fellow, Imperial College,1923; Lecturer in Zoology, Imperial College, 1926-1937, Reader, 1937-1964 and Professor of Zoology, 1964-1970, Imperial College, London; active in government service during the Second World War; Chief Rodent Officer, Ministry of Food, 1941-1945; Chairman, Advisory Committee, Infestation Control Laboratory, Ministry of Agriculture, Fisheries and Food, 1965-1974; Chairman, Farm Animals Welfare Committee, Ministry of Agriculture, Fisheries and Food, 1967-1974; Secretary-General of the International Congress of Zoology, 1959; died, 1974.
Publications: Practical Zoology. Instructions for dissection and preparation of elementary types of animals (Hutchinson, London, [1935]); XVth International Congress of Zoology, London 16-23 July 1958. Proceedings, edited jointly with N D Riley [London, 1958]; Grey Seals (Sunday Times, [London, 1962]).