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Born Titusville, Pennsylvania, 1912; educated Westminster School, 1925-1928; read Zoology at University College London, 1929-1933; MSc, University College London; PhD on locusts, including fieldwork in the Anglo-Egyptian Sudan, awarded 1939; Rockefeller Foundation Malaria Research Laboratory at Tirana, Albania, 1939; Research Officer in the Colonial Office's Middle East Anti-Locust Unit, 1942-1944, organising both Soviet and RAF crop-dusting aircraft; joined Vincent Brian Wigglesworth's Insect Physiology Unit at the Department of Zoology, Cambridge, 1946-1967; elected Fellow of the Royal Society, 1965; Deputy Chief Scientific Officer, Agricultural Research Council and Professor of Animal Behaviour, University of London at Silwood Park, 1967-1983; moved to the Department of Zoology, Oxford, 1983; research interests were aphids, moths and pheromones, insect flight patterns and from the later 1980s 'anthropomorphic misinterpretation of insect behaviour in various contexts'; Gold Medal (Zoology) of the Linnean Society, 1984; Wigglesworth Medal of the Royal Entomological Society, 1985; died, 1993.
Publications: A Conspectus of Aphids as Vectors of Plant Viruses with M F Day and Victor Frank Eastop (Commonwealth Institute of Entomology, London, 1962); Insect Polymorphism Editor (Royal Entomological Society, London, 1961); The new anthropomorphism (1992).