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Patrick Manson was born in 1844 and studied medicine at Aberdeen University, passing M.B. and C.M. in 1865. In 1866 he became medical officer of Formosa for the Chinese imperial maritime customs, moving to Amoy in 1871. Here, while working on elephantoid diseases, he discovered in the tissues of blood-sucking mosquitoes the developmental phase of filaria worms. From 1883 to 1889 he was based in Hong Kong, where he set up a school of medicine that developed into the university and medical school of Hong Kong. Returning to London, he became physician to the Seaman's Hospital in 1892. He played a central role in the development of tropical medicine as a distinct discipline, publishing on tropical diseases, being instrumental in the setting up of the London School of Tropical Medicine in 1899, and becoming physician and advisor to the Colonial Office in 1897. He propounded the theory that malaria was propagated by mosquitoes, a theory to be proved by Sir Ronald Ross (1857-1932). He was elected Fellow of the Royal Society in 1900 and awarded CMG, 1900, KCMG in 1903, and GCMG, 1912; he died in 1922.