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Born 1866; educated privately and King's College London; St Thomas's Hospital; University of Leipzig; Demonstrator in Biology and Physiology, King's College London and evening lecturer in Comparative Anatomy, 1887; Demonstrator in Physiology, University of Sydney, 1891; Professor of Physiology, University of Melbourne, 1901; 1903-1930 Director, Lister Insititute of Preventive Medicine, 1903-1930; Professor of Experimental Pathology, University of London; Chief of Division of Animal Nutrition of Australian Council for Scientific and Industrial Research, 1931-1933; Professor of Bio-chemistry and General Physiology, Adelaide University, 1931-1933; Committees and other appointments: served Gallipoli, Egypt, Palestine Expeditionary Force and France, 1915-1919; Lieutenant-Colonel, Australian Army Medical Corps and Consultant Pathologist, Australian Imperial Forces; Chairman of War Office Committee on anti-typhoid innoculation, 1904-1907; Member of Advisory Committee for Investigation of Plague in India, appointed jointly by the India Office, Royal Society and the Lister Institute, 1905-1913; Honours and awards: Fellow of the Royal College of Physicians, 1901; Fellow of the Royal Society, 1901; Companion of St Michael and St George, 1919; knighted, 1927; Doctor of Science (honorary) Sheffield and Dublin, Doctor of Laws (honorary) Edinburgh, Doctor of Civil Law (honorary) Durham, Master ofArts (honorary) Cambridge, Fellow of King's College London; Other information: married Edythe Cross, one daughter.