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Milne, son of Admiral Sir David Milne, was entered on the books of the Leander in 1817 but probably did not go to sea until 1820, when he joined the Conway on the South American Station. Again on this station between 1824 and 1830, he served in the ALBION, 1824 to 1825, the GANGES, 1825 to 1827, and the CADMUS, 1827 to 1830. He became a lieutenant in 1827. In 1837 he was promoted to commander into the SNAKE, North America and West Indies Station, where he operated against slavers, and in 1839 was appointed Captain of the CROCODILE on the same station. He transferred to the CLEOPATRA for a brief period in 1841 and then returned home. Milne was Flag-Captain to his father from 1842 to 1845 in the CALEDONIA, Devonport, and from 1845 to 1847 was in the St. Vincent at Portsmouth. He was on the Board of Admiralty until 1859, having become a rear-admiral in 1858. During the American Civil War Milne was Commander-in-Chief, North America and West Indies. On his return, he again joined the Board of Admiralty until 1869 when he commanded the Mediterranean Station for a year. In 1872 he was appointed Senior Naval Lord and after his retirement in 1876 he continued to be called upon for important tasks, including membership of the Carnarvon Commission on Colonial Defence, 1879 to 1882.