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Born, 1917; educated, Canford and Medical College of St Bartholomew's Hospital, -1943; House Surgeon, Senior House Surgeon and Chief Assistant to the Orthopaedic Unit, Hill End Hospital (St Bartholomew's Hospital), 1943-1946; Registrar to the Peripheral Nerve Injury Unit, Hill End Hospital, 1946-; demonstrator , Anatomy Department of the London School of Medicine for Women (now the Royal Free Hospital School of Medicine), 1946; Visiting Professor at Iowa State University; Reader, Royal Free Hospital School of Medicine; helped set up the Unit of Primatology at Royal Free Hospital; Director of the Primate Biology Program of the US National Museum, Smithsonian Institution; Director of a similar unit at Queen Elizabeth College, University of London; Visiting Professorship of Primate Biology at Birkbeck College, University of London; founder of the Primate Society of Great Britain; died, 1987.