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Born, 1903; Education: Emigrated to Canada with family in 1911; attended primary school in Calgary; Crescent Heights Collegiate Institute, Calgary; University of Alberta, Edmonton; Harvard (1924) and Harvard Gardens, Soledad, Cuba; Career: United Fruit Company, Columbia; US National Research Fellowship at Harvard; Institute of Entomology laboratory, Farnham Royal; research student at Gonville and Caius College, Cambridge (1931); Fellow of King's College, Cambridge (1933-2003); University Lecturer in Zoology, Cambridge University (1937-1965); war work for Ministry of Agriculture on the control of wireworms; Tutor for Advanced Students, King's College, Cambridge (1945-1951); collected speicimens in East Africa (1948-1949); field collection in Pakistan (1958-1959); Reader in Animal Ecology, University of Cambridge (1965-1971); Fellow of the Royal Society, 1956; died, 2003.