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Born in Great Berkhamsted, 1850; entered University College London, 1867; Demonstrator at University College London; Professor of Anatomy, University College London, 1877-1919; married Jenny Klingberg of Stockholm, god-daughter of the famous soprano Jenny Lind, 1884; three children, but his only son died young; examiner in anatomy at many universities, and to the Royal Colleges of Physicians and Surgeons; a founder member of the Anatomical Society of Great Britain and Ireland, and President, 1896-1897; knighted for his services to medical education in London and as inspector under the Vivi-Section Act (1876), 1919; Emeritus Professor of Anatomy, University College London; Fellow of the Royal College of Surgeons of England; LLD, Edinburgh; ScD, Dublin; Fellow of the Zoological Society; a member of various scientific societies overseas; died, 1930. Publications: with others, edited and contributed to Quain's Anatomy (9th and 10th editions) and Ellis's Demonstrations of Anatomy (10th and 11th editions).